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I. GENERAL INFORMATION
A. Methodology used to complete the review: In conducting this 5-year review, we relied on the best available information pertaining to historic and current distributions, life history, and habitats of this species. We announced initiation of this review and requested information in a published Federal Register notice with a 60-day comment period (74 FR 31972). We conducted an internet search, reviewed all information in our files, and solicited information from all knowledgeable individuals including those associated with academia and State conservation programs. Our sources include the final rule listing for this species under the Act; the recovery plan; peer reviewed scientific publications; unpublished field observations by Service, State and other experienced biologists; unpublished survey reports; and notes and communications from other qualified biologists or experts. The completed draft was sent to the other cooperating Service office and three peer reviewers for review. Comments were evaluated and incorporated into this final document as appropriate (see Appendix A). We did not receive any public comments during the 60-day open comment period.